1973-76 B.A., Oberlin College
1977-82 Ph.D., University of North Carolina, Chapel Hill
1982-85 Postdoctoral Research Associate, University of Massachusetts, Amherst 1984 Visiting Assistant Professor, Dartmouth College 1985-88 Assistant Professor, University of California, Irvine 1988-91 Associate Professor, University of California, Irvine 1991- Hannah Professor of Microbial Ecology, Michigan State University
1988 Committee of the Public Affairs Office of the Ecological Society of America on environmental applications of genetically engineered organisms 1988-1989 National Research Council Committee on Scientific Evaluation of the Introduction of Genetically Modified Microorganisms and Plants into the Environment 1989-1997 Associate Editor, Theoretical Population Biology 1990-1992 Associate Editor, Evolution 1990-1996 National Research Council Commission on Life Sciences 1990-1996 National Research Council Board on Biology 1991-1993 Editorial Board, Microbial Ecology 1991-1999 Editorial Advisory Board, Molecular Ecology 1994 National Science Foundation Population Biology Review Panel 1995 Vice-chair, Gordon Research Conference on "Population Biology and Evolution of Microbes and their Accessory Elements" 1995-1998 Editorial Board, The American Naturalist 1996-1998 Council Member, Society for the Study of Evolution 1997 Co-chair, National Research Council Workshop on "Ecology and Evolution of Infectious Diseases" 1999 Visiting Professor for Workshop on Experimental Evolution of Bacteria, Center for Population Biology, University of California, Davis 1999 Visiting Professor for Summer Workshop on Evolutionary Biology, University of Basel, Switzerland
1976 Phi Beta Kappa 1977 National Science Foundation Graduate Fellowship 1978 Sigma Xi 1986 American Society of Naturalists President's Award for Best Paper in The American Naturalist (with B. R. Levin) 1988 National Science Foundation Presidential Young Investigator 1992 John Simon Guggenheim Memorial Foundation Fellowship 1992 American Society of Naturalists President's Award for Best Paper in The American Naturalist (with M. R. Rose, S. C. Simpson and S. C. Tadler) 1996 John D. and Catherine T. MacArthur Foundation Fellowship 1997 Elected to Fellowship in American Academy of Microbiology 1998 Elected to Fellowship in American Academy of Arts and Sciences 1999 American Society for Microbiology, Division R Lectureship (Systematics and Evolution) 2006 Elected to Membership in National Academy of Sciences.